| 1951-1958 |
Wennington School, Wetherby, Yorkshire |
| 1959-1965 |
St. Catharine's College, Cambridge |
| 1961 |
Part I Natural Sciences Tripos: 1st class (Biochemistry, Botany, Organic Chemistry and Zoology) |
| 1962 |
Part II Natural Sciences Tripos: 1st class (Zoology) |
| 1962-1965 |
Agricultural Research Council. Studentship for research under Professor Sir Vincent Wigglesworth in Department of Zoology, Cambridge, ARC Unit of Insect Physiology |
| 1965 |
Ph.D., University of Cambridge |
| 1965-1966 |
Harkness (Commonwealth Fund) Fellowship for research and travel held at the Department of Biology, University of Virginia, Charlottesville, Va. (Head: Professor Dietrich Bodenstein) |
| 1966-1967 |
Harkness Fellowship held at Developmental Biology Center, Western Reserve University, Cleveland, Ohio (Head: Professor Howard Schneiderman) |
| 1967-1969 |
A.R.C. Postdoctoral Research Fellowship, Department of Genetics, University of Cambridge |
| 1969-1973 |
Open Research Fellowship, Caius College |
| 1969-2006 |
MRC Laboratory of Molecular Biology. Permanent member of scientific staff |
| 1976 |
Elected member of European Molecular Biology Organisation |
| 1977 |
Zoological Society of London's Medal |
| 1977-2010 |
Editor J. Embryology and experimental Morphology, became Development in 1987 |
| 1980 |
Dietrich Bodenstein Lecturer, University of Virginia |
| 1982 |
Walter Bauer Lecture, Helen Hay Whitney Meeting, New York
|
| 1983 |
Elected Fellow of the Royal Society |
| 1983-1989 |
Editorial Board of Cell |
| 1984-1986 |
Joint Head of Division of Cell Biology at Laboratory of Molecular Biology |
| 1987-1989 |
Editorial Board of EMBO Journal |
| 1994 |
De Camp Lecturer, Neurosciences Symposium, Columbia University, New York |
| 1994 |
Darwin Medal from Royal Society |
| 1996 |
Prize: "Vinci d'Excellence" Moet et Chandon, Paris |
| 1996 |
Inaugural Wigglesworth Memorial Lecture, XX International Congress of Entomology, Florence |
| 1996 |
Lecture to "Architectonics of Nature" Symposium. Princeton University 250th Anniversary celebrations |
| 1998 |
BBV Visiting Professor, Centro de Biología Molecular, Madrid |
| 2000 |
Waddington Medal from the British Society of Developmental Biology |
| 2000 |
Elected Fellow of the Royal Swedish Academy of Sciences |
| 2002 |
Visiting Miller Professor, University of California, Berkeley |
| 2004 |
Keynote Address, 45th Annual Drosophila Research Conference, Washington DC |
| 2006-2022 |
Director of Research, Department of Zoology, University of Cambridge |
| 2006-present |
MRC Laboratory of Molecular Biology. Emeritus Scientist |
| 2007 |
Prince of Asturias Award for Scientific and Technical Research, Oviedo (shared with Prof. Ginés Morata) |
| 2011 |
Lifetime Achievement Award from the Society for Developmental Biology (Bethesda, MD) |
| 2012 |
Honorary Fellowship of the Royal Entomological Society |
| 2013 |
Honorary Membership of the British Society of Developmental Biology |
| 2023-present |
Visitor of the Department of Zoology, University of Cambridge |